On Sunday, December 25, 2022 at 11:54:24 PM UTC-8,
[email protected] wrote:
On Sun, 25 Dec 2022 09:11:58 -0800 (PST), erik simpson
<[email protected]> wrote:
Title from https://www.tandfonline.com/doi/full/10.1080/03115518.2022.2132288
Abstract
A review of the Southern Hemisphere Mesozoic tribosphenic mammal fossil record supports the hypothesis that Tribosphenida arose in the Southern Hemisphere during the Early Jurassic, around 50 million years prior to the clade’s reliably dated first
appearance in the Northern Hemisphere. Mesozoic Southern Hemisphere tribosphenic mammals are known from Australia, Madagascar, South America and the Indian subcontinent, and are classified into three families: Bishopidae (fam. nov.), Ausktribosphenidae
and Henosferidae. These are stem therians, and considerable morphological evolution occurred within the lineage between the Jurassic and late Early Cretaceous. Important dental modifications include a graduated transition between premolars and molars,
development of molar wear facets V and VI, loss of facets for postdentary bones, reduction in the Meckelian groove and development of a true dentary angle. Previous classifications of Southern Hemisphere tribosphenic mammals are ambiguous
because information from the upper dentition has been lacking. Upper molars attributed to the late Early Cretaceous (Albian) Southern Hemisphere group Bishopidae fam. nov. are now known to possess a prominent protocone and stylar cusp C. We thus
consider bishopids to be the sister group to Theria.
This strongly suggests the origin of Tribosphenida was in the Jurassic, and >in the southern hemisphere. (The prevailing understanding is that the group was more or less synonymous with Boreosphenida (see https://en.wikipedia.org/wiki/Tribosphenida)
This is interesting to me. I understood mammal dentition to be a
defining characteristic of mammalian clades. But IIUC your cited
Wikipedia article, tribosphenic molars are found in several
paraphyletic groups. Are there other molar types which were once
considered diagnostic of a clade but are no longer?
I'm not that well-versed in the mammal fossil record of the Mesozoic. For a long time teeth
were the major part of that record, but my impression is that more recent finds have opened
things up some. Just glancing at phylogentic tree found by googling shows lots of polytomies
and short-branched clades. The overall view is pretty consistent, but there are lots of clades
that are less well-supported than you'd like.
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